[1] Univ Lisbon, Fac Med, Inst Farmacol & Neurociencias, P-1699 Lisbon, Portugal
[2] Inst Mol Med, Unidade Neurociencias, Lisbon, Portugal
[3] Univ Porto, Inst Biol Mol & Celular, Nerve Regenerat Grp, P-4100 Oporto, Portugal
[4] Univ Lisbon, Fac Farm, IMed UL, P-1699 Lisbon, Portugal