[1] Univ Shizuoka, Dept Biochem, Sch Pharmaceut Sci, Shizuoka, Japan
[2] JST, CREST, Tsukuba, Ibaraki, Japan
[3] COE Program 21st Century, Shizuoka, Japan
[4] Zhejiang Acad Med Sci, Hangzhou, Peoples R China
[5] Tokyo Metropolitan Inst Gerontol, Tokyo, Japan
[6] Nagoya City Univ, Dept Biol Struct & Biomol Engn, Grad Sch Pharmaceut Sci, Nagoya, Aichi, Japan