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Guizhou Karst Environmental Ecosystems Observation and Research Station, Ministry of Education, Key Laboratory of Karst Georesources and Environment, Ministry of Education, College of Resources and Environmental Engineering, Guizhou University, Guiyang,550 Guizhou Karst Environmental Ecosystems Observation and Research Station, Ministry of Education, Key Laboratory of Karst Georesources and Environment, Ministry of Education, College of Resources and Environmental Engineering, Guizhou University, Guiyang,550