[1] Univ Estadual Paulista UNESP, Dept Ciencias Biol, LEDA, FCL, Ave Dom Antonio 2100,Parque Univ, Assis, SP, Brazil
[2] Univ Estadual Paulista UNESP, Inst Biociencias, Dept Zool, Rua Prof Dr Antonio Celso Wagner Zanin 250, Botucatu, SP, Brazil
[3] Univ Sao Paulo, Inst Biociencias, Dept Zool, Rua Matao,Travessa 14,101 Cidade Univ, Sao Paulo, Brazil
[4] Univ Sao Paulo, Ctr Biol Marinha CEBIMar, Rodovia Manoel Hypolito Rego,Km 131-50, Sao Sebastiao, SP, Brazil