[1] Univ Rennes, CNRS, Geosci Rennes, UMR 6118, F-35000 Rennes, France
[2] Univ Paris, Univ Paris Diderot, UFR Sci Vivant, F-75013 Paris, France
[3] EPHE, Inst Systemat Evolut Biodiversite ISYEB, Museum Natl Hist Nat, CNRS,SU,UA, 57 Rue Cuvier, F-75231 Paris 05, France
[4] Chinese Acad Sci, State Key Lab Palaeobiol & Stratig, Nanjing Inst Geol & Palaeontol, Nanjing 210008, Peoples R China
[5] Chinese Acad Sci, Ctr Excellence Life & Paleoenvironm, Nanjing 210008, Peoples R China
[6] Univ Bonn, Inst Geowissensch, Bonn 53115, Germany