[1] Chinese Acad Fishery Sci, South China Sea Fisheries Res Inst, Guangzhou 510300, Guangdong, Peoples R China
[2] Guangdong Prov Key Lab Fishery Ecol & Environm, Guangzhou 510300, Guangdong, Peoples R China
[3] State Ocean Adm, Inst Oceanog 3, Lab Marine Biol & Ecol, Xiamen 361005, Peoples R China
[4] Collaborat Innovat Ctr Deep Sea Biol, Hangzhou 310012, Zhejiang, Peoples R China