[1] Ehime Univ, Grad Sch Agr, 3-5-7 Tarumi, Matsuyama, Ehime 7908566, Japan
[2] Ehime Univ, Ctr Adv Technol Environm, Grad Sch Agr, 3-5-7 Tarumi, Matsuyama, Ehime 7908566, Japan
[3] Australian Museum, Res Inst, 1 William St, Sydney, NSW 2010, Australia
[4] Ehime Univ, Fac Agr, 3-5-7 Tarumi, Matsuyama, Ehime 7908566, Japan