New combinations in South American Glandularia (Verbenaceae)

Morphological and karyological features support the transfer of four species from Verbena L. to Glandularia J. F. Gmelin: G. balansae (Briquet) N. O'Leary, G. paraguariensis (Moldenke) N. O'Leary, G. tecticaulis (Troncoso) N. O'Leary, and G. thymoides (Chamisso) N. O'Leary. Nevertheless, this group has features, such as brief style length and the funnelform and short corolla tubes, that are typical of Verbena. These four species represent a group informally named Balansae that is interesting from an evolutionary point of view. Two leetotypifications are made for G. balansae and G. thymoides.