[1] Lebanese Univ, Earth & Life Sci Dept, Fac Sci 2, POB 26110217, Fanar, Matn, Lebanon
[2] Tiroler Landesmuseum Ferdinandeum, Josef Schraffl Str 2a, A-6020 Innsbruck, Austria
[3] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, State Key Lab Palaeobiol & Stratig, Nanjing 210008, Peoples R China
[4] Chinese Acad Sci, Ctr Excellence Life & Paleoenvironm, Nanjing 210008, Peoples R China