[1] SE Asian Fisheries Dev Ctr, Dept Aquaculture, Iloilo 5021, Philippines
[2] Florida Inst Technol, Dept Biol Sci, Fish Ecophysiol Lab, Melbourne, FL 32901 USA
[3] Sch Environm Resources & Dev, Aquaculture & Aquat Resources Management, Asian Inst Technol, Pathum Thani, Thailand
[4] Univ Philippines, Inst Biol, Coll Sci, Quezon City 1101, Philippines
[5] Fisheries Res Agcy, Kanagawa, Japan