[1] Univ Witwatersrand, Inst Human Evolut, ZA-2050 Johannesburg, South Africa
[2] Univ Witwatersrand, Sch Geosci, Bernard Price Inst Palaeontol Res, ZA-2050 Johannesburg, South Africa
[3] Univ Bordeaux 1, Inst Prehist & Geol Quaternaire, CNRS, UMR 5199, F-33405 Talence, France
[4] George Washington Univ, Dept Anthropol, Washington, DC USA
[5] Univ Witwatersrand, Sch Geog Archaeol & Environm Studies, ZA-2050 Johannesburg, South Africa