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Laboratory of Industrial Microbiology and the Laboratory of Fermentation Physiology and, Applied Microbiology, Division of Applied Life Sciences, Graduate School of Agriculture, Kyoto University, Kitashirakawa-oiwakecho, Sakyo-ku, Kyoto 606-8502, Japan Laboratory of Industrial Microbiology and the Laboratory of Fermentation Physiology and, Applied Microbiology, Division of Applied Life Sciences, Graduate School of Agriculture, Kyoto University, Kitashirakawa-oiwakecho, Sakyo-ku, Kyoto 606-8502, Japan