[1] Inst Nacl Mata Atlantica INMA, BR-29650000 Santa Teresa, ES, Brazil
[2] Inst Marcos Daniel, Projeto Bromeligenous, Av Eugenio Pacheco Queiros S-N, BR-29090160 Vitoria, ES, Brazil
[3] Univ Fed Espirito Santo UFES, Programa Posgrad Biol Anim, Vitoria, ES, Brazil
[4] Consejo Nacl Invest Cient & Tecn, Museo Argentino Ciencias Nat Bernardino Rivadavia, Div Herpetol, Angel Gallardo 470,C1405DJR, Buenos Aires, DF, Argentina
[5] Univ Estado Minas Gerais, Dept Ciencias Biol, Campus Uba, BR-36502000 Uba, MG, Brazil