[1] Southwest Univ, Key Lab Freshwater Fish Reprod & Dev, Minist Educ, Chongqing 400715, Peoples R China
[2] Southwest Forestry Univ, Key Lab Conserving Wildlife Small Populat Yunnan, Kunming 650224, Yunnan, Peoples R China
[3] Michigan State Univ, E Lansing, MI 48825 USA
[4] Rutgers Univ Newark, Boyden Hall,195 Univ Ave, Newark, NJ 07102 USA
[5] Honghe Univ, Coll Life Sci & Technol, Mengzi 661199, Yunnan, Peoples R China