[1] Zvitambo Inst Maternal & Child Hlth Res, Harare, Zimbabwe
[2] Univ Liverpool, Inst Infect Vet & Ecol Sci, Liverpool, England
[3] Queen Mary Univ London, Blizard Inst, London, England
[4] Queen Mary Univ London, Blizard Inst, 4 Newark St, London E1 2AT, England