[1] MNHN, Dept Milieux & Peuplements Aquat, UMR CNRS Biol Organ Marins & Ecosyst 5178, F-75231 Paris 05, France
[2] Univ Gdansk, Inst Oceanog, Dept Marine Ecosyst Functioning, Lab Estuarine Ecol, PL-81378 Gdynia, Poland
[3] Univ La Rochelle, IFREMER, CNRS, CRELA UMR 6217, F-17000 La Rochelle 1, France