[1] Univ Leeds, Genet & Cell Biol, Leeds, W Yorkshire, England
[2] Univ Leeds, Fac Biol Sci, Dept Mol & Cell Biol, Leeds LS2 9JT, W Yorkshire, England
[3] Univ Leeds, Fac Biol Sci, Astbury Ctr Struct Mol Biol, Leeds LS2 9JT, W Yorkshire, England
[4] Univ Helsinki, Quantitat Biol, Helsinki, Finland
[5] Univ Helsinki, Helsinki Inst Life Sci HiLIFE, Helsinki 00014, Finland
[6] Univ Helsinki, Fac Biol & Environm Sci, Helsinki 00014, Finland