[1] Univ New South Wales, Evolut & Ecol Res Ctr, Sydney, NSW 2052, Australia
[2] Univ New South Wales, Sch Biol Earth & Environm Sci, Sydney, NSW 2052, Australia
[3] Sydney Inst Marine Sci, Chowder Bay Rd, Mosman, NSW 2088, Australia
[4] Port Stephens Fisheries Inst, Locked Bag 1, Nelson Bay, NSW 2315, Australia