[1] Kyushu Univ, Grad Sch Pharmaceut Sci, Dept Life Innovat, Fukuoka, Fukuoka, Japan
[2] Kyushu Univ, Grad Sch Pharmaceut Sci, Dept Mol & Syst Pharmacol, Fukuoka, Fukuoka, Japan
[3] Univ Toyama, Grad Sch Med & Pharmaceut Sci, Dept Appl Pharmacol, Toyama, Japan
[4] Kyushu Univ, Grad Sch Med, Dept Dermatol, Fukuoka, Fukuoka, Japan
[5] Johns Hopkins Univ, Sch Med, Solomon H Snyder Dept Neurosci, Baltimore, MD USA
[6] Johns Hopkins Univ, Sch Med, Ctr Sensory Biol, Baltimore, MD USA