[1] Univ Manchester, Arthrit Res UK Ctr Genet & Genom, Manchester M13 9PL, Lancs, England
[2] Cent Manchester Fdn Trust, NIHR Manchester Musculoskeletal Biomed Res Unit, London, England
[3] Univ Manchester, Manchester Acad Hlth Sci, Manchester M13 9PL, Lancs, England
[4] Univ Erlangen Nurnberg, Human Genet, D-91054 Erlangen, Germany
[5] Univ Crete, Mol Med & Human Genet & Internal Med, Iraklion, Greece
[6] Univ Leeds, NIHR Leeds Musculoskeletal Biomed Res Unit, Leeds Inst Rheumat & Musculoskeletal Med, Leeds LS2 9JT, W Yorkshire, England
[7] Meath Hosp, Med Clin, Mol Med, Dublin, Ireland
[8] Meath Hosp, Dublin, Ireland
[9] Univ Dublin Trinity Coll, Dublin 2, Ireland
[10] Univ Bath, Royal Natl Hosp Rheumat Dis & Pharm & Pharmacol, Bath BA2 7AY, Avon, England
[11] Univ Roma Tor Vergata, Biomed & Prevent, Rome, Italy
[12] Fdn Santa Lucia IRCCS, Lab Mol Genet UILDM, Rome, Italy
[13] Keele Univ, Sci & Technol Med, Rheumatol, Haywood Hosp,Hlth Serv Res Unit, Keele, Staffs, England
[14] Univ Coll Dublin, St Vincents Univ Hosp, UCD Sch Med & Med Sci, Rheumatol, Dublin, Ireland
[15] Univ Coll Dublin, Conway Inst Biomol & Biomed Res, Dublin, Ireland
[16] Goethe Univ Frankfurt, Grp Translat Med & Pharmacol, Rheumatol & Fraunhofer IME Project, D-60054 Frankfurt, Germany
[17] Cent Manchester Fdn Trust, Kellgren Ctr Rheumatol, NIHR Manchester Biomed Res Ctr, Manchester, Lancs, England
[18] CSIC, Parasitol & Biomed Lopez Neyra, Granada, Spain
[19] IIS IPrincesa, Rheumatol, Hosp La Princesa, Madrid, Spain