[1] Hong Kong Polytech Univ, Dept Land Surveying & Geoinformat, Hong Kong 999077, Peoples R China
[2] Univ Hong Kong, Fac Sci, Sch Biol Sci, Hong Kong 999077, Peoples R China
[3] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol & Biodivers, 16 Lincui Rd, Beijing 100101, Peoples R China
[4] Beijing Normal Univ, Fac Geog Sci, State Key Lab Earth Surface Proc & Resource Ecol, Beijing 100875, Peoples R China