[1] Univ Sistan & Baluchestan, Dept Biol, Fac Sci, Zahedan, Iran
[2] Shiraz Univ, Dept Biol, Coll Sci, Ichthyol Res Lab, Shiraz, Iran
[3] Univ Munich, Dept Earth & Environm Sci, Palaeontol & Geobiol, Munich, Germany
[4] Univ Munich, GeoBio Ctr LMU, Munich, Germany
[5] Ferdowsi Univ Mashhad, Dept Biol, Fac Sci, Azadi Sq,POB 9177948974, Mashhad, Khorasan Razavi, Iran
[6] Ferdowsi Univ Mashhad, Inst Appl Zool, Fac Sci, Res Dept Zool Innovat, Mashhad, Iran