[1] Chinese Acad Sci, Inst Zool, Key Lab Anim Ecol & Conservat Biol, Beijing, Peoples R China
[2] Univ Chinese Acad Sci, Int Coll, Beijing, Peoples R China
[3] Yunnan Univ, Inst Int Rivers & Ecosecur, Kunming, Yunnan, Peoples R China
[4] Chinese Acad Sci, Chengdu Inst Biol, Chengdu, Sichuan, Peoples R China
[5] Wildlife Conservat Div, Sichuan Forestry Dept, Chengdu, Sichuan, Peoples R China