[1] Shimane Univ, Estuary Res Ctr, Matsue, Shimane, Japan
[2] Japan Fisheries Res & Educ Agcy, Fisheries Resources Inst, Demersal Fish Resources Div, Fisheries Stock Assessment Ctr, Niigata, Niigata, Japan
[3] Japan Fisheries Res & Educ Agcy, Socioecol Syst Div, Fisheries Resources Inst, Yokohama, Kanagawa, Japan
[4] Hokkaido Univ, Grad Sch Environm Sci, Hakodate, Hokkaido, Japan
[5] Hokkaido Univ, Fac Fisheries Sci, Hakodate, Hokkaido, Japan
[6] Japan Fisheries Res & Educ Agcy, Fisheries Resources Inst, Highly Migratory Resources Div, Fisheries Stock Assessment Ctr, Yokohama, Kanagawa, Japan
[7] Fisheries & Oceans Hakodate, Hakodate, Hokkaido, Japan
[8] Hokkaido Univ, Field Sci Ctr Northern Biosphere, Hakodate, Hokkaido, Japan
[9] Japan Fisheries Res & Educ Agcy, Coastal & Inland Fisheries Ecosyst Div, Fisheries Technol Inst, Nagasaki, Japan
[10] Japan Fisheries Res & Educ Agcy, Marine Fisheries Res & Dev Ctr, Natl Res & Dev Agcy, Yokohama, Kanagawa, Japan
[11] Japanese Inst Fisheries Infrastruct & Communities, Chiyoda Ku, Tokyo, Japan