[1] Univ Zurich, Inst Evolutionary Biol & Environm Studies, Zurich, Switzerland
[2] Univ Oxford, Dept Zool, Oxford OX1 3PS, England
[3] McGill Univ, Dept Biol, Montreal, PQ H3A 1B1, Canada
[4] Univ Minnesota, Dept Forest Resources, St Paul, MN USA
[5] Univ Wageningen & Res Ctr, Nat Conservat & Plant Ecol Grp, Wageningen, Netherlands
[6] Swiss Ornithol Inst, Ctr Populat Anal, Sempach, Switzerland