[1] Fabis Consulting Ltd, Chestnut Lane, Nottingham NG11 0AE, England
[2] MTA Ctr Ecol Res, Klebelsberg K U 3, H-8237 Tihany, Hungary
[3] Museum Natl Hist Nat, European Top Ctr Biol Divers, 57 Rue Cuvier, FR-75231 Paris 05, France
[4] European Commiss, Joint Res Ctr, Via E Fermi 2749, I-21027 Ispra, VA, Italy
[5] Norwegian Inst Nat Res NINA Trondheim, POB 5685 Torgard, N-7485 Trondheim, Norway