[1] Hunan Normal Univ, Coll Life Sci, Changsha 410081, Hunan, Peoples R China
[2] Hubei Univ, Sch Life Sci, State Key Lab Biocatalysis & Enzyme Engn, 368 Youyi Rd, Wuhan 430062, Hubei, Peoples R China
[3] Hubei Univ, Sch Life Sci, Ctr Behav Ecol & Evolut, 368 Youyi Rd, Wuhan 430062, Hubei, Peoples R China
[4] Natl Univ Singapore, Dept Biol Sci, 14 Sci Dr 4, Singapore 117543, Singapore