[1] Chinese Acad Sci, South China Sea Inst Oceanol, State Key Lab Trop Oceanog, Guangzhou 510301, Peoples R China
[2] Southern Marine Sci & Engn Guangdong Lab Guangzho, Guangzhou 511458, Peoples R China
[3] Chinese Acad Sci, Innovat Acad South China Sea Ecol & Environm Engn, Guangzhou 510301, Peoples R China
[4] Chinese Acad Sci, Daya Bay Marine Biol Res Stn, Shenzhen 518121, Peoples R China
[5] Guangdong Technion Israel Inst Technol, Environm Sci & Engn Program, 241 Daxue Rd, Shantou 515063, Guangdong, Peoples R China
[6] Univ Chinese Acad Sci, Coll Earth & Planetary Sci, Beijing 100049, Peoples R China