[1] Arthritis Res UK Ctr Genet & Genom, Manchester, Lancs, England
[2] Ctr Epidemiol, Manchester, Lancs, England
[3] Univ Manchester, Arthritis Res UK Ctr Genet & Gen, Manchester, Lancs, England
[4] Cent Manchester Fdn Trust, NIHR Manchester Musculoskeletal BRU, Manchester, Lancs, England
[5] Univ Manchester, Manchester Acad Hlth Sci Ctr, Manchester, Lancs, England
[6] Newcastle Univ, Inst Cellular Med, Newcastle Upon Tyne, Tyne & Wear, England
[7] NIHR Leeds Musculoskeletal Biomed Res Unit, Leeds, W Yorkshire, England
[8] Manchester Acad Hlth Sci Ctr, Cent Manchester NHS Fdn Trust, Musculoskeletal Biomed Res Ctr, Natl Inst Hlth Res Manchester, Manchester, Lancs, England
[9] Univ Manchester, Ctr Musculoskeletal Res, Manchester Acad Hlth Sci Ctr, Arthritis Res UK,Ctr Genet & Gen, Manchester, Lancs, England
[10] Univ Coll Dublin, Conway Inst, UCD Sch Med & Med Sci, Dublin, Ireland