[1] Russian Acad Sci, Caspian Inst Biol Resources Dagestan Sci Ctr, M Gadzhiev Str 45, Makhachkala 367000, Russia
[2] Dagestan State Univ, M Gadzhiev Str 43a, Makhachkala 367000, Russia
[3] Russian Acad Sci, Inst Zool, Univ Skaya Emb 1, St Petersburg 199034, Russia
[4] Sorbonne Univ, MNHN, UPMC, EPHE,CNRS, 57 Rue Cuvier,CP 50,Entomol,45 Rue Buffon, F-75005 Paris, France