[1] Univ Bordeaux, UMR CNRS EPOC 5805, Allee Geoffroy St Hilaire Bat 18N, F-33615 Pessac, France
[2] PSL Univ, EPHE, Allee Geoffroy St Hilaire Bat 18N, F-33615 Pessac, France
[3] Univ Bern, Inst Plant Sci, Altenbergrain 21, CH-3013 Bern, Switzerland
[4] Univ Bern, Oeschger Ctr Climate Change Res, Altenbergrain 21, CH-3013 Bern, Switzerland
[5] Swiss Fed Inst Forest Snow & Landscape Res WSL, A Ramel 18, CH-6593 Cadenazzo, Switzerland