[1] Chinese Acad Sci, Yantai Inst Coastal Zone Res, Yantai 264003, Peoples R China
[2] Chinese Acad Sci, Yantai Inst Coastal Zone Res YIC, CAS Key Lab Coastal Environm Proc & Ecol Remediat, Yantai 264003, Peoples R China
[3] YICCAS, Shandong Key Lab Coastal Environm Proc, Yantai 264003, Peoples R China
[4] Chinese Acad Sci, Ctr Ocean Mega Sci, 7 Nanhai Rd, Qingdao 266071, Peoples R China
[5] Chinese Acad Sci, Inst Geog Sci & Nat Resources Res, Key Lab Ecosyst Network Observat & Modeling, Beijing 100101, Peoples R China
[6] Univ Chinese Acad Sci, Coll Resources & Environm, Beijing 100190, Peoples R China
[7] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[8] Ludong Univ, Sch Resources & Environm Engn, Yantai 264039, Shandong, Peoples R China
[9] Jiangxi Normal Univ, Sch Geog & Environm, Nanchang 330022, Jiangxi, Peoples R China
[10] Beijing Forestry Univ, Sch Ecol & Nat Conservat, Beijing 100083, Peoples R China
[11] Wageningen Univ & Res, Land Use Planning Grp, NL-6700 HB Wageningen, Netherlands
[12] Beijing Normal Univ, Coll Life Sci, Minist Educ, Key Lab Biodivers Sci & Ecol Engn, Beijing 100875, Peoples R China