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Univ Autonoma Barcelona, Fac Biociencies, Systemat & Evolut Vasc Plants UAB, Associated Unit,CSIC,Dept Biol Anim Vegetal & Eco, ES-08193 Bellaterra, Spain Univ Autonoma Barcelona, Fac Biociencies, Systemat & Evolut Vasc Plants UAB, Associated Unit,CSIC,Dept Biol Anim Vegetal & Eco, ES-08193 Bellaterra, Spain