[1] Ctr Rech Halieut, UMR EME 212, Inst Rech Dev, F-34203 Sete, France
[2] Univ Perpignan, CRIOBE, CNRS, EPHE,USR 3278, F-98729 Moorea, France
[3] Univ Perpignan, Ctr Biol & Ecol Trop & Mediterraneenne, CNRS, EPHE,UPVD,UMR 5244, F-66860 Perpignan, France
[4] Univ Perpignan, Ctr Biol & Ecol Trop & Mediterraneenne, UR128, IRD, F-66860 Perpignan, France
[5] Univ Antilles Guyane, Equipe DYNECAR, Marine Biol Lab, F-97159 Pointe A Pitre, Guadeloupe, France
[6] Univ Paris 06, LOCEAN, CNRS, IRD,MNHN,UMR 7159, F-75252 Paris 05, France
[7] Ctr Rech Halieut Mediterraneenne & Trop, UMI IRD 209, F-34203 Sete, France
[8] Ctr Rech Halieut Mediterraneenne & Trop, UPMC, UMMISCO, F-34203 Sete, France
[9] UMR UdS CNRS 7178, Dept Ecol Physiol & Ethol, Inst Pluridisciplinaire Hubert Curien, F-67087 Strasbourg, France