[1] Ovidius Univ Constanta, Fac Nat & Agr Sci, Al Univ 1, Constanta 900470, Romania
[2] Univ Tecn Particular Loja, Dept Ciencias Nat, Calle Marcelino Champagnat S-N, Loja, Ecuador
[3] Univ Nacl Loja, CITIAB, Ciudadela Univ, EC-110101 La Argelia, Loja, Ecuador
[4] Univ Liege, Behav Biol Unit, Lab Fish & Amphibian Ethol, 22 Quai van Beneden, B-4020 Liege, Belgium
[5] Pontificia Univ Catolica Ecuador, Dept Ciencias Biol, Museo Zool, Ave 12 Octubre 1076 & Roca,Apartado 17-01-218, Quito, Ecuador