[1] Inst Natl Biodiversidad, Unidad Invest, Casilla 17-07-8976, Quito, Ecuador
[2] Univ Nacl Loja, Loja, Ecuador
[3] USFQ, Lab Zool Terr, Colegio Ciencias Biol & Ambientales, Quito 170901, Ecuador
[4] USFQ, Colegio Ciencias Biol & Ambientales, Museo Zool, Quito 170901, Ecuador
[5] Pontificia Univ Catolica Ecuador, Ecuela Biol, Museo Zool QCAZ, Quito, Ecuador
[6] Kings Coll London, Dept Geog, London, England
[7] Minist Ambiente, Direcc Nacl Biodiversidad, Quito, Ecuador