[1] Univ Calif San Diego, Med, San Diego, CA 92103 USA
[2] NHS Greater Glasgow & Clyde, Virol, Glasgow, Lanark, Scotland
[3] NHS Greater Glasgow & Clyde, Med, Glasgow, Lanark, Scotland
[4] NHS Greater Glasgow & Clyde, Brownlee Ctr Infect Dis, Glasgow, Lanark, Scotland
[5] Glasgow Caledonian Univ, Sch Hlth & Life Sci, Glasgow, Lanark, Scotland
[6] NHS Greater Glasgow & Clyde, Publ Hlth, Glasgow, Lanark, Scotland
[7] Univ Edinburgh, Inst Evolutionary Biol, Edinburgh, Midlothian, Scotland