[1] Tokyo Metropolitan Univ, Dept Biol Sci, Hachioji, Tokyo 1920397, Japan
[2] Res Inst Evolutionary Biol Inc, Setagaya Ku, Tokyo 1580098, Japan
[3] Univ Tokyo, Univ Museum, Bunkyo Ku, Tokyo 1130033, Japan
[4] Utah State Univ, Dept Biol, Logan, UT 84322 USA
[5] Nagasaki Univ, Inst Trop Med, Dept Int Hlth & Med Anthropol, Nagasaki 8528523, Japan
[6] SUNY Syracuse, Coll Environm Sci & Forestry, Dept Environm & Forest Biol, Syracuse, NY 13210 USA
[7] Chiba Univ, Marine Biosyst Res Ctr, Uchiura, Chiba 2995502, Japan
[8] Univ Liege, Dept Funct & Evolutionary Entomol, Gembloux Agrobio Tech, B-5030 Gembloux, Belgium