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Chinese Acad Sci, Wuhan Inst Virol, Mol Virol & Viral Immunol Res Grp, State Key Lab Virol, Wuhan 430071, Hubei, Peoples R China Chinese Acad Sci, Wuhan Inst Virol, Mol Virol & Viral Immunol Res Grp, State Key Lab Virol, Wuhan 430071, Hubei, Peoples R China