[1] Univ Fed Rio Grande do Sul, Inst Biociencias, Dept Bot, Programa Posgrad Bot, Campus Vale, BR-91509900 Porto Alegre, RS, Brazil
[2] Univ Fed Santa Catarina, Ctr Ciencias Biol, Dept Bot, Programa Posgrad Biol Fungos Algas & Plantas, Campus Reitor Joao David Ferreira Lima, BR-88040900 Florianopolis, SC, Brazil
[3] Univ Utah, Nat Hist Museum Utah, Salt Lake City, UT USA
[4] Univ Utah, Dept Biol, Salt Lake City, UT 84112 USA
[5] Royal Ontario Museum, Dept Nat Hist, Toronto, ON, Canada
[6] Univ Toronto, Dept Ecol & Evolutionary Biol, Toronto, ON, Canada