[1] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Tibetan Environm Changes & Land Surface P, Bldg 3,Courtyard 16,Lin Cui Rd, Beijing 100101, Peoples R China
[2] Univ Chinese Acad Sci, 19 A Yuquan Rd, Beijing 100049, Peoples R China
[3] CAS Ctr Excellence Tibetan Plateau Earth Sci, Bldg 3,Courtyard 16,Lin Cui Rd, Beijing 100101, Peoples R China
[4] Lincoln Univ, Dept Agr & Environm Sci, 820 Chestnut St, Jefferson City, MO 65101 USA
[5] Lincoln Univ, Cooperat Res Program, 820 Chestnut St, Jefferson City, MO 65101 USA