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Minist Agr & Rural Affairs, Coll Anim Sci & Technol, Shandong Prov Key Lab Anim Biotechnol & Dis Contro, Key Lab Efficient Utilizat Non Grain Feed Resource, Tai An 271018, Shandong, Peoples R China Minist Agr & Rural Affairs, Coll Anim Sci & Technol, Shandong Prov Key Lab Anim Biotechnol & Dis Contro, Key Lab Efficient Utilizat Non Grain Feed Resource, Tai An 271018, Shandong, Peoples R China