[1] Swiss Ctr Faunal Cartog, CH-2000 Neuchatel, Switzerland
[2] Univ Lausanne, Inst Ecol, CH-1015 Lausanne, Switzerland
[3] Utah State Univ, USGS Biol Resources, Utah Cooperat Fish & Wildlife Res Unit, Logan, UT 84322 USA
[4] Stanford Univ, Dept Stat, Stanford, CA 94305 USA