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Japan Fisheries Res & Educ Agcy, Hokkaido Natl Fisheries Res Inst, Toyohira Ku, 2-2 Nakanoshima, Sapporo, Hokkaido 0620922, Japan
Japan Fisheries Res & Educ Agcy, Tohoku Natl Fisheries Res Inst, 4-9-1 Sakiyama, Miyako, Iwate 0270097, Japan Japan Fisheries Res & Educ Agcy, Hokkaido Natl Fisheries Res Inst, Toyohira Ku, 2-2 Nakanoshima, Sapporo, Hokkaido 0620922, Japan