[1] Miyazaki Univ, Miyazaki Med Coll, Dept Pharmacol, Miyazaki 8891692, Japan
[2] Dokkyo Med Univ, Dept Pharmacol, Mibu, Tochigi 3210293, Japan
[3] Univ Occup & Env Hlth, Sch Med, Dept Pharmacol, Yahatanishi Ku, Kitakyushu, Fukuoka 8078555, Japan
[4] Fukuoka Univ, Fac Med, Dept Pharmacol, Jhonan Ku, Fukuoka 8140180, Japan
[5] Oita Univ, Fac Med, Dept Pharmaceut Med Commun, Hasama, Yufu 8795593, Japan