[1] Univ Amsterdam, Inst Biodivers & Ecosyst Dynam, Ctr Geoecol Res, NL-1098 SM Amsterdam, Netherlands
[2] Univ Nottingham, Sch Geog, Nottingham NG7 2RD, England
[3] Univ Essen Gesamthsch, Inst Geog, D-45117 Essen, Germany
[4] Univ Nacl Colombia, ICN, Lab Palinol & Paleoecol, Bogota 7495, Colombia