[1] Univ South Dakota, Dept Biol, Vermillion, SD 57069 USA
[2] Univ Quebec Rimouski, Dept Biol Chim & Geog, Rimouski, PQ, Canada
[3] Ctr Sci Biodiversite Quebec, Grp Rech Environm Nordiques BOREAS, Ctr Etud Nordiques, Rimouski, PQ, Canada
[4] Univ Pretoria, DST NRF Ctr Excellence, Dept Zool & Entomol, Percy FitzPatrick Inst, Hatfield, South Africa
[5] South African Natl Biodivers Inst, South African Res Chair Conservat Physiol, Pretoria, South Africa
[6] Lund Univ, Dept Biol, Lund, Sweden