[1] Fac Sci Tunis, Dept Biol, Lab Microorganismes & Biomol Act, Tunis 2092, Tunisia
[2] CBBC, Lab Physiol Mol Plantes, Hammam Lif 2050, Tunisia
[3] Inst Natl Rech Genie Rural Eaux & Forets, Lab Ecol & Ameliorat Sylvo Pastorale, Ariana 2080, Tunisia
[4] Inst Natl Rech Agron Tunisie, Lab Protect Vegetaux, Ariana 2080, Tunisia
[5] Inst Super Educ & Format Continue, Lab Mat, Le Bardo 2019, Tunisia