[1] Aichi Gakuin Univ, Dept Oral Anat, Sch Dent, Nagoya, Aichi, Japan
[2] Harvard Univ, Dept Human Evolutionary Biol, Cambridge, MA 02138 USA
[3] Wellesley Coll, Dept Biol Sci, Wellesley, MA 02181 USA
[4] Wake Forest Univ, Dept Anthropol, Winston Salem, NC 27109 USA
[5] Kyoto Univ, Grad Sch Sci, Kyoto 6068501, Japan
[6] Kyoto Univ, Primate Res Inst, Kyoto 6068501, Japan