[1] Chinese Acad Sci, Northwest Inst Plateau Biol, Key Lab Adaptat & Evolut Plateau Biota, Xining 810008, Peoples R China
[2] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol & Biodivers, Beijing 100085, Peoples R China
[3] CAS Ctr Excellence Tibetan Plateau Earth Sci, Beijing 100101, Peoples R China
[4] Univ Chinese Acad Sci, Beijing 100049, Peoples R China